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Biological Sciences Courses

Students must confirm their registration with their instructors by the second class meeting, or their registration may be canceled. All Common Core courses and some advanced courses have laboratories; in the following course descriptions, L indicates courses with a laboratory. No more than four courses below Biological Sciences 193 may be taken without consent of the senior adviser. Students who have taken or are taking courses in Biological Sciences Sequence 1 may not enroll in other 100-level courses (except 195) without consent of the senior adviser.

Common Core Sequences

Sequence 1. This sequence is an intensive introduction to the biological sciences. It meets the same Common Core objectives as other sequences; however, its courses are more advanced and are designed to prepare students for 200-level courses in the biological sciences. Past experience suggests that if you meet the prerequisites, these courses will not be too difficult. This sequence is open to all who meet the prerequisites, regardless of future concentration, but is meant for students who are considering a concentration in the biological sciences. A year of college chemistry, two quarters of calculus, or the equivalents are prerequisites for the latter three courses.

Students ordinarily begin Sequence 1 (193 or 195) in their first year, and these courses are without prerequisites. Students may complete their Common Core requirement with either 193-194-196, 195-196-197, or 196-197-198.

Sequence 2. Students in Sequences 2-4 choose one from a group of selected courses for each of three quarters, beginning with the autumn quarter. The biological sciences are concerned with understanding the natural universe at many different levels of organization and complexity, from molecules and cells to populations, species, and ecosystems. Autumn quarter courses in Sequence 2 focus on the biology of cells and give students an appreciation of the cellular and molecular mechanisms that underlie higher levels of biological organization. Winter quarter courses focus on the biology of organisms, and consider how these cellular and molecular mechanisms are integrated and regulated in multicellular organisms. Spring quarter courses consider the evolutionary processes by which organisms and species have arisen.

Sequence 3. The courses in Sequence 3 are designed to achieve the same educational goals as do those in Sequence 2, but they present biological concepts in a different order. Autumn quarter courses focus on organisms and consider various features that characterize organismal function. Winter quarter courses then emphasize the genetic mechanisms by which information is transmitted from parents to their offspring. In the spring quarter, students can take a course in evolutionary biology from Sequence 2, or choose a course that emphasizes the process and progress of science.

Sequence 4. Sequence 4 presents the diversity of the biological sciences in a different order from Sequences 2 or 3. Autumn quarter courses introduce students to evolutionary and ecological processes. Winter quarter courses are the same courses as in Sequence 3, and emphasize genetic mechanisms. Finally, spring quarter courses illustrate how evolutionary processes and genetic mechanisms contribute to the functioning of cells and organisms, or focus on the process of scientific discovery.

Sequence 5 examines knowledge and values in biology as they influence and are influenced by cultural ideas and human values of the nineteenth and twentieth centuries. It emphasizes critical reading and evaluation of original texts. This sequence is ideal for students who want a general overview of biology or wish to sample a great many different topics.

Sequence 6 examines the basic principles of modern biology and their application to larger societal issues, including human disease and the environment. It is intended for students who do not plan to concentrate in the biological sciences. The general emphasis is on an appreciation of the current impact of the biotechnological revolution and its ramifications for the future in the context of a liberal education.

Sequences 7 and 8. There are two six-quarter sequences in the natural sciences. Ordinarily students must complete an entire natural sciences sequence to fulfill the Common Core biological sciences requirement and may not enter these sequences in the middle. As described below, however, students may take the last three courses in the Natural Sciences 151-156 sequence as Biological Sciences 154-156 (Sequence 9) without having taken Natural Sciences 151-153. For descriptions of these courses, see the Natural Sciences section of the catalog.

Sequence 9 is the latter half of the natural sciences sequence "Form and Function in the Natural World." Students may join the sequence in either autumn or winter quarter, and must complete all three courses in this sequence. Biological Sciences 154 requires an elementary knowledge of chemistry or the consent of the instructor.

Sequence 10 provides an overall introduction to the biological sciences from an organismal viewpoint. It is intended principally for students who have strong quantitative skills and are considering careers as research scientists or physicians. It emphasizes areas of the biological sciences that currently provide exciting research opportunities and are amenable to quantitative or physiochemical analysis. Both lectures and laboratories are taught by faculty, who are also available for discussions of potential research opportunities. Laboratory exercises are specially designed to complement the lectures and involve both data collection and mathematical and computer modeling of biological systems.

Admission to the sequence is by consent of instructor. Students should be prepared to develop the mathematical skills necessary to solve biological problems requiring the use of differential equations as well as linear algebra.

Sequence 11 uses the theme of growth and development as a focus around which to organize the presentation of many of the exciting advances of contemporary biology. The courses in this sequence emphasize cellular aspects of growth and development, but pay attention to the molecular mechanisms that underlie these cellular processes and the ways in which the integration of cellular processes is essential for the survival of organisms.

Sequence 12 stresses fundamental concepts of cell biology, genetics, and development. The courses in this sequence stress the importance of these basic biological mechanisms in the lives of organisms and the evolutionary conservation of cellular structures and developmental processes.

Common Core Courses

102. Introductory Genetics. PQ: Autumn quarter course in Sequence 3 or 4 and high school science/chemistry. The basic principles of genetics are presented in terms of gene theory as it developed from transmission and molecular investigations of eukaryotic and prokaryotic species. E. Garber. Winter. L.

103. Introduction to Evolution. PQ: High school science. The first part of the course presents the data of patterns and phenomena lending support to evolutionary theory: the fossil record, diversity of organisms, and modern approaches to phylogenetic reconstruction. The second part discusses the genetic basis of evolution and the third covers processes and mechanisms of evolution: natural selection and speciation. The material is presented in the form of a scientific inquiry, posing data, developing hypotheses, and demonstrating appropriate forms of testing. Three lab exercises and a visit to the scientific collections at the Field Museum allow hands-on experience of data collecting and data analysis. Two discussions explore the impact of evolutionary theory on our society. P. Sierwald. Autumn. L.

104. Human Heredity. PQ: BioSci 135. This course examines the principles of molecular and general genetics, emphasizing the possible application of recent findings to humans and society. Discussion section required. B. Strauss. Winter. L.

106. Biotic Crises. PQ: Winter quarter course in Sequence 2 or 3. In this course, crises are defined as major instabilities that occur with sufficiently low frequency so that species cannot adapt to or compensate for them. Rates and patterns of crises and extinctions are important aspects of the evolution and organization of ecological systems and can take place in relatively short time frames (ecological time) or in longer ones (evolutionary time). The disasters can be physical or biological in origin (for example, flooding or a viral epidemic). Waves of extinctions in the geological past have been recently interpreted as due to meteoritic (or cometary) impacts. We ask a basic question: Are the cumulative effects of rare phenomena more significant biologically than "normal" phenomena? M. Nitecki. Spring. L.

107. Evolution. PQ: High school biology. An introduction to evolutionary biology, including discussion of population genetics, natural selection, adaptation, speciation, origin of novelties, and evolution above the species level, using examples from both living and extinct organisms. The last third of the course concentrates on the history of life with special attention to why major changes in the flora and fauna have taken place. J. Masterson. Autumn. L.

108. Essentials of Cell Biology. PQ: High school science/chemistry. This course surveys biochemical aspects of cellular structure and function. Topics include protein synthesis and secretion, endocytosis and receptor-mediated uptake, storage, energy transformations, biochemical regulation, and cell division. These biochemical processes in the cell are related to the health and disease of the organism of which the cell is an integral part. P. Soltys. Autumn. L.

109. Biology of Cells and Tissues. PQ: High school science/chemistry. The organization of living matter can be analyzed at many levels. Cells are the most basic level for self-sustaining life. We discuss the structure and organization of cells (with emphasis on the composition of cells) and how cells synthesize, organize, and utilize matter. Topics include DNA structure and replication; protein structure and synthesis; the importance of carbohydrates and lipids for cellular function; and cellular respiration, reproduction, and communication. Discussion section required. M. Musch. Autumn. L.

110. Bacteria, the Biosphere, and Biotechnology. PQ: Autumn quarter course in Sequence 3 or 4. The course serves as an introduction to general principles of biology by studying the smallest living cells, bacteria, and their roles in the environment, symbiosis, disease, and biotechnology. The fundamentals of biological function are illustrated by emphasis on the biosynthesis and action of macromolecules, the operation of cellular control circuits, the role of environmental sensing, and new insights into the importance of intercellular communication. The importance of both natural and synthetic genetic engineering are discussed with regard to issues of wider social concern, such as antibiotic control of infectious disease and biotechnological application of bacteria. J. Shapiro. Winter. L.

111. Evolution. PQ: Winter quarter course in Sequence 2 or 3. This course examines the pattern and process of evolution across many biological levels, from the gene, to the individual, to groups of individuals, on up to species and beyond. Students learn how ecology, genetics, and molecular and developmental biology, as well as paleontology, all come together in the study of evolution. The course covers specific topics in evolution, including natural selection and adaptation, molecular evolution and selfish DNA, the evolution of sex, the evolution of behavior, speciation, and the origin of diversity. Labs include exposure to the Field Museum's collections and exhibits. R. Bieler. Spring. L.

112. Developmental Biology. PQ: BioSci 104. The purpose of this course is to study the processes of development from one cell, the fertilized egg, to a whole organism. The emphasis is on understanding the concepts of development. Topics include fertilization, cell lineage, the derivation of tissues and organs, and cell and tissue interactions. Basic genetic principles essential for the understanding of development are also examined. M. Moscona. Spring. L.

115. Fundamentals of Cell Biology. PQ: High school science. This course represents an overview of the basic molecular processes that determine cellular structure and function. Topics include synthesis of biological macromolecules, organelle structure and energy flow, and molecular aspects of intercellular communication. Nucleic acid biochemistry, gene regulation, and recombinant DNA technology are also discussed. Special emphasis is placed on mechanisms which regulate cell division and differentiation in normal organismal development and in disease states, such as cancer. P. Marone. Autumn. L.

117. Introduction to Physiology. PQ: High school science/chemistry. This course is an introduction to the study of physiology and includes topics such as digestion, absorption, and transport of foodstuffs; the kidney and homeostasis; the physiology of bone; the reproductive system; the cardiovascular system; neurobiology; and endocrinology. Evening discussion section required. H. Rochman. Autumn, Winter. L.

118. Human Physiology in Health and Disease. PQ: Autumn quarter course in Sequence 2. Physiology is the study of biological function. In this course, we begin to explore the physiology of the human organism, including such functions as digestion, reproduction, circulation, sensation, and movement. By comparing human function in health with the derangements of select disease processes, we endeavor to gain a deeper understanding of vital activities each of us engages in daily. Along the way, we also consider the historical foundations of our understanding of human physiology and ponder the distinctively human significance of our vegetative and animal functions. R. Gunderman. Winter. L.

119. The Hungry Earth: Light, Energy, and Subsistence. PQ: Autumn quarter course in Sequence 4 and winter quarter course in Sequence 3 or 4. The pressure exerted on the resources of the earth by the growing human population makes a broad knowledge of biology, especially plant biology, relevant to finding possible solutions to crucial problems society is facing. In this context, we discuss the principles of energy conversion by plants and their diversity, growth, and reproduction. The origin of domesticated plants, their improvement by genetic engineering, the interaction of plants with pathogens, and the adaptation of organisms to the rotation of the earth are also examined. M. Ruddat. Spring. L.

120. Brain, Perception, and Behavior. PQ: Autumn quarter course in Sequence 2 and high school chemistry. This course provides a survey of contemporary brain research, emphasizing the neural bases of perception and motor control. Topics include the ionic basis of spike generation and axonal conduction; synaptic transmission; neurotransmitters and drugs; language and the brain; functional neurophysiology of the visual system; the physiological basis of learning and memory; and the neural control of movement. Phylogenetic comparisons, nervous system development, philosophical issues, and analogies between brains and computers are discussed as appropriate. H. Fozzard, Staff. Winter. L.

121. Nutrition in the Health of Populations. PQ: Winter quarter course in Sequence 3 or 4 and high school science. This course examines human nutritional needs from a population perspective, the relationship between food choices and human health, and how scientific information translates into recommendations on nutrition. Students learn how to assess the validity of scientific research that provides the basis for advice about how to eat healthfully. Topics include nutritional assessment, nutritional epidemiology, factors affecting the nutritive value of foods, nutrification of foods, nutrition fraud, and nutrition policy and goals. Class projects are designed to help students apply their learning by critiquing nutritional health claims and current nutrition policy issues. P. Strieleman. Spring. L.

122. Plant Structure and Function. This course incorporates an integrated approach to the design and function of plants, including physiological, morphological, and anatomical perspectives. Photosynthesis, water transport, adaptation of plants to extreme environments, biotechnology, sexual reproduction in plants, and other topics are covered. S. Mayer. Autumn. L.

123. Biology of Motion. This course explores all kinds of motion in organisms, from the streaming movements within a cell to the migratory flights of hawks. Topics include cellular locomotion, muscle contraction, biological pumps, the use of antagonistic muscles to move hard or soft skeletons, burrowing, swimming, and walking. The cellular and metabolic mechanisms of motility are related to the physiology, behavior, and ecological context of an organism. We also see how energetics, scale, and evolutionary history constrain the mechanisms an organism uses for motility. Students participate in hands-on investigations in class, such as measuring the skeletons of a human and of other animals to compare their locomotory adaptations. T. Colton. Autumn. L.

124. Perspectives in Health. PQ: Students must attend the first class to confirm enrollment. No exceptions. This course utilizes an investigative approach to examine scientific concepts related to health and disease. Case studies provide the basis for the evaluation of information and the promotion of decision making regarding health issues. Diverse topics are explored, including nutrition, sexual health, fertility, and chronic and communicable diseases. M. H. Maskay. Autumn. L.

125. Heredity: Cells and Genes (How Does Heredity Work?). PQ: High school biology; high school chemistry helpful. An introduction to molecular and cell biology, with an emphasis on the central role of genetic processes in the maintenance, growth, and differentiated functions of cells. Beginning with analysis of Mendel's paper and subsequent discoveries about the chromosomal basis of inheritance, we end with results of recombinant DNA technology. Discussion section required. J. Spofford. Autumn. L.

127. The Science of Nutrition. PQ: Autumn quarter course in Sequence 2 and high school science. Students must attend the first class to confirm enrollment. No exceptions. This course promotes an understanding of the principles of human nutrition by exploring current controversies. The underlying biological mechanisms of nutrient utilization are investigated in relation to the role of diet and nutrition in health and disease. M. H. Maskay. Winter. L.

131. The Workings of the Human Brain: A Disease-Oriented Approach. PQ: Autumn quarter course in Sequence 4 and winter quarter course in Sequence 3 or 4. Neurological deficits from stroke and other neurological diseases offer the opportunity to learn about the functions of the human brain. In this course, a brief review of the structure of the human brain is followed by descriptions of individuals with neurological deficits. Images from magnetic resonance imaging (MRI) and neurophysiological recordings are shown. A modern approach to localizing brain function, functional MRI, is discussed. The validity of the methods used to probe brain functions and problems of interpretation of the data are addressed. Through this exercise and selected readings, the student receives an overview of brain function and an appreciation of neurological disorders. A. Noronha. Spring. L.

133. Animal Behavior: Function and Evolution. PQ: Winter quarter course in Sequence 3 or 4. Concepts of animal behavior are studied with emphasis on experimental approaches to understanding particular behaviors. Topics include the physiological basis of behavior, the nature of territoriality, sexual selection and reproductive strategies, social interactions, and the extent to which behavior is genetically determined. The emphasis is on hypothesis testing as a means of comprehending behavioral patterns. Topics are examined within an evolutionary context. L. Houck. Spring. L.

135. Molecular Conservation of Cell Function. Basic processes occurring in living cells have been conserved from bacteria to plants to humans. This course discusses the organization of parts of the cell and stresses the molecular conservation of both structure and function in diverse groups of organisms. T. Martin, K. Chiang. Autumn. L.

136. Conservation of Developmental Mechanisms in Biology. PQ: BioSci 104. This course explores the extreme diversity of developmental patterns for embryonic development and indicate how recent molecular information has established an unexpected level of molecular conservation. J. Austin. Spring. L.

137. Introduction to Cell Biology. PQ: Autumn quarter course in Sequence 4 and winter quarter course in Sequence 3 or 4; high school science/chemistry. This course is intended to give students an awareness of recent advances in our understanding of the working of the cell. The structure and function of the cell and its organelles are described. Mechanisms for controlling cell number, growth, and survival are presented. Particular emphasis is given to cell-cell and cell-microenvironment communication. Molecules, signaling devices, and pathways involved in transmitting information from the outside in and from the inside out of the cell are discussed, taking examples from the nervous and immune systems. S. Szuchet. Spring.

138. Reproductive Biology. PQ: Autumn quarter courses in Sequence 4 and winter quarter course in Sequence 3 or 4. This course reviews the mammalian reproductive system and its functions, considering developmental aspects and including comparative reproductive biology of lower animals. The biology of human reproduction is featured and some reference is made to dysfunction, reproductive failure, and fertility regulation. Topics include gametogenesis, gametes and gamete transport, ovulation, fertilization, egg transport, early embryonic development, implantation, and pregnancy. The course stresses the biochemistry of male and female sex hormones, their regulation, and the significance of their action for the synchronization of biological and biochemical events in reproduction. G. F. B. Schumacher. Spring. L.

139-140-141. Growth and Development: I Molecular Aspects, II Cellular Aspects, III Integrative Aspects. PQ: Course must be taken in sequence; high school biology and chemistry. This sequence covers the evolving concepts of molecular, cellular, and organismal biology. These interrelated courses focus on some of the major advances in modern biology that address the fundamental molecular processes leading to normal and abnormal cell growth. The autumn course focuses on the proteins and genes that contribute to cell growth, using the reproductive system as a model. The winter course considers the molecular aspects of cellular interaction and communication. The spring course considers how these processes are integrated in response to infection and injury, using the immune system as a model. G. Greene, Staff, Autumn; M. Rosner, Staff, Winter; F. Fitch, Staff, Spring. L.

143. Current Topics in Genetics: From Mendel to Genetic Engineering. PQ: Autumn quarter course in Sequence 3 or 4. A topical approach is used to explore past and current scientific developments and trends in genetics. Such topics as Mendelian genetics, the history and practical implications of the discovery of DNA molecular structure, organellar DNA and its evolutionary significance, recombinant DNA and genetic engineering, the detection of human genetic diseases and their potential treatment with gene therapy, and DNA fingerprinting are introduced and discussed. These topics are used to develop an understanding of approaches to scientific problem solving and to explore the social implications of recent genetic advances. Class discussions of controversial issues in genetics are an important component of the course. Discussion section required. J. Feder. Winter. L.

145. Trying on Your Genes: Topics in Genetics. PQ: Autumn quarter course in Sequence 3 or 4. Students are introduced to the study of genetics as an exciting and challenging branch of modern biology. The overall goal of this course is for students to gain understanding of how geneticists approach biological questions. By studying current, newsworthy issues in genetics and molecular biology, students learn the basic concepts of genetics, heredity, and recombinant DNA technology. Topics include how mutations give rise to cancer, how genes program cells to differentiate, and the genetic basis of disease. For each example, emphasis is placed on understanding the strategy used to address the problem. B. Berg. Winter. L.

146. Human Genetics. PQ: Autumn quarter course in Sequence 3 or 4. The objective of this course is to provide students with an understanding of the basic principles of human genetics and how they are applied to the study of human disease and the delivery of medical genetic services. The course reviews the methods for estimating recurrence risks for genetic diseases, applying recombinant DNA technology to the diagnosis of genetic diseases, providing genetic counseling to families at risk for genetic diseases, and developing screening and diagnostic genetics programs in the community. The ethical implications of the potential for preventing and treating human diseases using new technological advancements are also considered. Discussion section required. C. Ober. Winter. L. Not offered 1995-96; will be offered 1996-97.

147. Genetics in an Evolutionary Perspective. PQ: Autumn quarter course in Sequence 3 or 4 and high school advanced algebra and science/chemistry. The course covers the historical development of theories of heredity and evolution, beginning prior to Darwin and Mendel and culminating in the "synthetic theory" of evolution, in which macroevolutionary processes are explained using models from population genetics. Major current disputes concerning evolutionary theory and genetics and their historical applications in biology and the social sciences are discussed. A lab using computer simulations illustrates the major processes of evolution and genetics, the characteristics of some of the simpler models in population biology, and the strategy and tactics of mathematical model building. W. Wimsatt. Winter. L.

148. Regulation of Human Physiological Systems. PQ: Autumn quarter course in Sequence 2. Open only to students not expecting to receive their degree in June 1996. This course examines the manner by which the cells that constitute a multicellular organism such as a human being are maintained with a relatively stable chemical and thermal environment despite wide variations in the external environment in which the organism lives. The focus is on three organ systems (and their interrelationships) that are crucial to this achievement: the circulatory system, the kidneys, and the neuroendocrine system. The class format emphasizes active class participation. Attendance at each class session required. P. Hoffmann. Winter. L.

149. Genetics. PQ: Autumn quarter course in Sequence 3 or 4. This course examines the general principles of genetics and shows how they operate at the organismal and molecular levels. For each, examples center on human-based issues. Our aim is not to examine each area exhaustively but rather to present a broad overview, concentrating on the research and problem-solving strategies employed by workers in these fields. To experience this process in the classroom, we use strategic simulations. In the final portion of the course, we build our own models to study how simulations can in themselves be very powerful and interesting tools. In this way, we gain insight into the phenomena we are trying to model and see how models can guide explanation, interpretation, and discovery in science. Discussion section required. J. Kruper. Winter. L.

151. The Biological Revolution and Its Implications. This course is intended for students not planning advanced work in the biomedical sciences but is open to all. It examines the biological bases and technology of molecular biology and genetic engineering, considers present and future outcomes of scientific progress in this area, and discusses the implications of such progress. M. Feder. Autumn. L.

152. Ecology. This course examines interactions between organisms and their biotic/abiotic environment, and the reasons populations vary over space and time. Topics include climate, major biomes of the world, and physical factors affecting distribution of species; biogeochemical cycles and energy flow in ecosystems; interactions among organisms, such as predation, competition, and symbiosis; factors that limit populations; the structure and development of ecological communities; and the ecological impact of human activities. Some topics are covered through case studies, in which students are exposed to methods of research and encouraged to discuss and evaluate controversial issues from an ecological point of view. Field trips to museums, zoos, and field sites required. E. Fielding. Autumn. L.

154. Living Cells (=NatSci 154). PQ: Elementary chemistry or consent of instructor. This course considers the basic attributes of living cells with special emphasis on how the structure of macromolecules determines their functional role in cellular processes. Emphasis is also placed on the nature of inquiry and discovery in modern molecular biology and on providing sufficient basic knowledge and vocabulary to enable a nonprofessional to follow and appreciate the reporting of future developments and their ramifications. E. Goldwasser. Autumn. L.

155. Multicellular Organisms (=NatSci 155). PQ: BioSci 154 or consent of instructor. This course provides a description of the relationships between structure and function in a multicellular organism. Examples are chosen that illustrate how the demands of a particular function determine its structure, and how the existence of certain structures in the developmental or evolutionary history of an organism affects its function. Examples are drawn from a variety of animals, including the human organism. L. Straus. Winter. L.

156. Individuals, Populations, and Groups (=NatSci 156). PQ: BioSci 155 or consent of instructor. This course focuses on biological processes at and above the individual level: variation, selection, and the evolution of adaptations; resources and hazards in natural environments; evolution of life history patterns; aggregations and social groups; structure, growth, regulation, and conservation of natural populations and communities; and political and ethical aspects of human population problems. Vertebrate animals will be emphasized. P. Soltys. Spring. L.

157. The Diversity of Life through Time. PQ: Winter quarter course in Sequence 2 or 3. This course surveys the major systematic groups of plants and animals and examines biological diversity through time (including the current biological diversity/extinction crisis). Topics include the nature and classification of species; processes of speciation and growth; aspects of morphology and body plans; and ecological specializations such as plant pollination, animal/plant coevolution, and feeding and locomotion. Evolution and biogeography of organisms and biotas are considered within a broad context of biological and physical influences. M. Westneat, T. Lammers. Spring. L.

160. Animal Reproductive Strategies. Successful reproduction is essential to the survival and evolution of species. This course examines the great variety of reproductive strategies that organisms use to propagate their genes and consider why such diversity exists. Topics covered include the evolution of sexual reproduction, mate identification and courtship behavior, sexual selection, and parental care; examples are drawn from a wide variety of species, including humans. Reproductive physiology is introduced as needed. Practical applications to captivate management and conservation of endangered species are discussed. L. Hornig. Autumn. L.

162. Evolution and Human Diversity. PQ: Winter quarter course in Sequence 2 or 3. This course examines evolution and human diversity from before Darwin to the present from a biological, historical, ethical, and societal perspective. Evolution, gradual or sudden, species extinction, and the role of natural selection, migration, and genetic drift in human diversity are analyzed first. The biology, ideology, and politics of race, and the perennial fascination with eugenics of societies from the early Greeks to the present are used as models of the use and abuse of genetics. The evolution of skin color and population variation in blood groups, enzymes, hemoglobins, chromosomes, Tay-Sachs disease, and cystic fibrosis are then categorized from a population and clinical perspective, with forays into DNA polymorphisms, forensics, consanguinity, human and ape homology, and evolution today. Finally, the economic, social, and political aftermath of genetic technology on the individual, the family, and society serves as a paradigm for the unpredictable outcome of scientific discovery. J. Bowman. Spring. L.

166. Medicine and Human Understanding. PQ: Winter quarter course in Sequence 3 or 4. Medicine may be described as applied human biology. As such, it moves back and forth between general theories about health and disease and personal contact with individual human beings. It tests our capacities to direct and apply science in the service of human welfare and provides poignant case studies of larger questions about human virtue and suffering. In this course, our readings interweave biological, philosophical, and literary approaches to the science and art of healing. R. Gunderman. Spring. L.

167. Biology of Organisms I. PQ: Previous or concurrent registration in a first-year calculus sequence. This course considers the diversity of animal body plans from an evolutionary and developmental viewpoint. Lectures cover major groups of animals, systematic relationships between animals as determined by cladistic methods, and the relationship between developmental and evolutionary processes. R. Lombard, M. Martindale. Autumn. L.

168. Biology of Organisms II. PQ: BioSci 167. The second course in this sequence emphasizes neurobiology and biomechanics. Lectures cover the integrative properties of neurons as determined by their passive electrical properties and voltage-gated conductances, the properties of groups of neurons, the biomechanics of muscles and bones as related to terrestrial locomotion, and the application of fluid mechanics to understanding the movements of animals in air and water. P. Ulinski, M. Dickinson. Winter. L.

169. Biology of Organisms III. PQ: BioSci 168. The third course in this sequence considers the relationships of organisms to their environments and to each other. It deals with the environmental problems posed by aquatic and terrestrial environments, the genetic and biochemical mechanisms involved in environmental adaptations, and social behavior. D. Crawford, D. Margoliash. Spring. L.

172. Introduction to Genetics. PQ: Autumn quarter course in Sequence 3 or 4 and high school science/chemistry. This course is an introduction to the basic concepts in genetics, beginning with Mendelian inheritance and ending with current topics of investigation in the field. Emphasis is on the molecular aspects of gene expression, regulation, and modification. P. Soltys. Winter. L.

174. Physiology of Stress. PQ: Autumn quarter course in Sequence 2. Principles of physiological regulatory mechanisms are explained by analyzing the response of animals, including humans, to stress--exposure to environmental extremes, work overload (mental and physical), injury, infection, malnutrition, and so on. Appropriate areas of organ physiology, neurobiology, cell biology, biochemistry, and ultrastructures are covered. R. Zak. Winter. L.

175. Introduction to Evolutionary Biology. PQ: Winter quarter course in Sequence 2 or 3 and high school science. The course introduces students to major facets of evolutionary biology, from the historical development of evolutionary ideas to modern aspects of molecular evolution. The first part presents the database of patterns lending support to evolutionary theory: the fossil record, diversity of organisms, and modern aspects of their classification and distribution. The second part discusses the genetic basis and molecular evolution. The third part covers mechanisms of natural selection, adaptation, and speciation. Four laboratories and a visit to the scientific collections at the Field Museum introduce students to data collection and analysis in the light of evolutionary theory. Two discussion periods explore the impact of evolutionary theory on our society. P. Sierwald. Spring. L.

176-177-178. Biology and Society I, II, III. PQ: Courses must be taken in sequence. This sequence is designed to develop the liberal arts of critical thinking and critical writing, with biology as their subject. Students examine knowledge and values in biology and how they are integrated with the cultural ideas and human values of the nineteenth and twentieth centuries. To understand the methods and ideas at work in biological science, we read textbooks and original papers: ecology, cell biology, and genetics in autumn; human physiology of the reproductive and nervous systems in winter; and evolutionary biology in spring. Twice each week, we meet for interactive lectures. In the textbooks we find the positive content of biology, and in the original papers we find major decision points in the history of ideas that are crucial in the intellectual effort of the Western world. Once each week, we meet for small group discussions that focus on contemporary issues in society of concern in the biology under consideration that quarter: environmental ethics, genetic screening, manipulation, and therapy in autumn; biomedical ethics and reproductive technologies, neural transplants into human brains, brain-mind-selfhood, and artificial intelligence in winter; and Darwin and literature, philosophy and religion, the race-IQ debate, sociological critiques of sociobiology, and evolutionary ethics in spring. This course emphasizes careful preparation of texts and active participation in each class. R. Holmes. Autumn, Winter, Spring. L.

182. Immunology: Basic Concepts and Its Role in Health and Disease. PQ: Autumn quarter course in Sequence 4 and winter quarter course in Sequence 3 or 4. Recommended for premedical students. This course deals with the fundamentals of immunology, with particular reference to immunology as it relates to defense against infection and to immunologic processes in certain disease states. General principles of cellular and humoral immunity are covered in detail with appropriate clinical correlations. Other topics include the complement system, phagocytosis, metabolic processes relating to bacterial killing, and antiviral defense systems. Allergic phenomena and autoimmune processes are discussed as examples of immunologically mediated disease. J. Nachman. Spring. L.

185-186-187. Biological Basis of Health and Disease I, II, III. PQ: Courses must be taken in sequence; high school science, biology, and chemistry helpful. This three-quarter sequence covers the evolving concepts of molecular, cellular, and organismal biology through the study of normal and abnormal states in health and disease. These interrelated courses focus on some of the major advances in modern biology and medicine that have contributed to our understanding of the origins and maintenance of life and explore the fundamental mechanisms by which illness develops and may be treated or prevented. The process of scientific investigation leading to these discoveries is considered from a historical perspective, with emphasis on the impact of these discoveries on society and on the delivery of health care. Discussion section optional. Staff. Autumn, Winter, Spring. L.

193-194. Ecology, Genetics, and Evolution. This two-quarter sequence surveys the major principles of ecology, Mendelian genetics, and evolutionary biology. Topics in ecology include demography and life histories, competition, predation, and the interspecific interactions that shape the structure of ecological communities. The fundamentals of classic transmission genetics, the major experimental studies in heredity, and the consequences of Mendelian inheritance for evolution are presented. We discuss in a quantitative way the constellation of evolutionary forces that shapes adaptation and the diversity of all biological systems, including mutation, random genetic drift, and natural selection. Specific topics in this section of the course include sexual selection and the evolution of sex dimorphism as well as the evolution of social behaviors. The overall goals of this two-quarter sequence are (1) to teach students how to think about the population processes that affect the numbers and genetic diversity of living systems and (2) to expose students to current problems in ecology and evolution suitable for advanced study in upper-level courses or in lab rotations. Discussion section required. M. Wade, T. Wootton. Winter, Spring. L.

195. Biological Diversity. This course examines the diversity of living things, including viruses, bacteria, protistans, fungi, plants, and animals, as well as the diversity of the environments in which they occur and interact. The intent is to provide a firm grounding in adaptations and life processes as they occur in organisms in their natural setting, and to set the stage for a consideration of the major principles and processes underlying the function of living things and their evolution. Although this course is open to any undergraduate, it is an intensive introduction to biological diversity designed for those considering a concentration in the biological sciences. Staff. Spring. L.

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196. Cell and Molecular Biology. PQ: Chem 111-112-113 and two quarters of calculus, or equivalent. An introduction to molecular and cellular biology, emphasizing the unity of biochemical processes in cells. The following topics are covered: the structure, function, and synthesis of nucleic acids and protein; cell structure; functional organization; intermediary metabolism; energetics; some properties of differentiated cells; aspects of chemical evolution and the origins of cellular structure; and genetic engineering. Discussion section required. L. Mets. Autumn. L.

197. Genetics. PQ: BioSci 196. Fundamentals of genetic mechanisms as they apply to simple transmission genetics and developmental biology. The course stresses concepts and their implications for understanding the inheritance of simple traits and for discovering the molecular basis for developmental processes, with special emphasis on the embryonic development of model organisms. B. Strauss, A. Mahowald. Winter. L.

198. Evolutionary Biology. PQ: BioSci 196-197. This course surveys the major principles of evolutionary biology. Topics include the evidence for evolution, the history of life, the mechanisms of evolution (mutation, selection, and genetic drift), adaptation, speciation, the origin of evolutionary novelties, the origin of life, and human evolution. Lectures are combined with lab and discussion classes. B. Charlesworth, J. Coyne. Spring. L.

Advanced-Level Courses

There are three types of advanced courses. General courses are courses in which instructors present the general principles and recent developments for broad areas within the biological sciences. Such courses are usually offered on a regular basis, either annually or biennially. Specialized courses either focus on a topic of particular interest to the instructor or examine topics at a more advanced level than in general courses. Such courses are offered less regularly, as warranted by student and faculty interest. Unless otherwise stated, most general and specialized biological sciences courses assume mastery of the material in Biological Sciences 195-196-197-198 or its equivalent. These courses are open to all students in the College who satisfy the prerequisites. Courses on ethical and societal implications of the biological sciences are, as the name implies, of interest to all students in the College. Such courses generally either have no prerequisite or have any Common Core biology sequence as a prerequisite.

The following table is intended to aid students in planning programs of courses. Course offerings may change from year to year. The letters in parentheses indicate 1995-96 courses that satisfy the developmental biology (d); organismal biology (o); and ecology, populations, and behavior (e) requirements of the concentration program.

General Courses

199. Introduction to Research. PQ: Consent of a faculty sponsor and the Undergraduate Research and Honors Committee. Students are required to submit the College Reading and Research Course Form. This course is graded P/F. May be elected for up to three quarters. This course cannot be used to satisfy any requirements for the biological sciences concentration. Staff. Summer, Autumn, Winter, Spring. L.

200. Introduction to Biochemistry. PQ: BioSci 196-197; Chem 217-218 or 220-221-222 recommended. This course fulfills the biochemistry requirement for the biological sciences concentration. The chemical nature of cellular components, enzymes, and mechanisms of enzyme activity, energy interconversions, and biosynthetic reactions, including template-dependent processes and some aspects of control mechanisms, are studied. H. Friedmann. Autumn, Spring. L.

203. Introduction to Biophysics and Biophysical Chemistry. PQ: Chem 217-218 or 220-221, or consent of instructor. An introductory course emphasizing concepts of physical chemistry important in the interactions of biological macromolecules, with emphasis on dynamics and kinetics. The course focuses on basic aspects of secondary and tertiary structure, the origin and basis of electrostatic and hydrophobic interactions, and dynamical properties of proteins. The importance of concepts of diffusion and transport in biological processes is also treated. Problem sets are coordinated with lectures. M. W. Makinen, R. Astumian. Spring.

207. Cell Biology. PQ: BioSci 200 or equivalent. This course surveys gene organization and expression; functions of the cell nucleus, cytoskeleton, and cytoplasmic structures; and cell-cell interactions and signaling. E. Taylor, Autumn. T. Martin, J. Miller, Winter.

208. Immunobiology. PQ: Chem 111-112-113 or equivalent and Common Core biology. This course presents an integrated coverage of the tactics and logistics of immune phenomena and conveys the elegance of the biological solutions evolved by multicellular organisms in their fights against infectious agents. Immune phenomena are presented as unique evolutionary adaptations of vertebrates operating in the context of ancillary defense mechanisms. The various types of countermeasures evolved by pathogens are also discussed, with particular emphasis on HIV and discussions on AIDS. Discussion section required. J. Quintans. Winter. L.

209. Immunobiology. PQ: Chem 111-112-113 or equivalent and Common Core biology. This course is identical to BioSci 208 except that it does not have a lab. Discussion section required. J. Quintans. Autumn.

212. Neurobiology. PQ: Common Core biology and consent of instructor. This is a first advanced course in neurobiology. Topics include cellular properties of neurons and glia (their structure and function); the membrane potential, action potentials, and mechanisms of synaptic transmission; the organization of neurons into neuronal pathways; and sensory transduction. The lab focuses primarily on electrophysiological techniques used in the analysis of issues fundamental to neural processing at the cellular level, including monitoring membrane potential, carrying out voltage clamp of native and cloned ion channels, and investigating the control of synaptic transmission. D. Hanck. Spring. L.

213. Neurobiology. PQ: Common Core biology and consent of instructor. This course is identical to BioSci 212 except that it does not have a lab. D. Hanck. Spring.

215. Experimental Bacterial Genetics: Genome Alignment in Rhodobacter. PQ: BioSci 200 and consent of instructor. This course satisfies two of the lab requirements (cell biology and genetics) for the biological sciences concentration. This course is intended to introduce students to current methods of investigating the molecular genetics of bacteria. Students construct cosmid and phage libraries, screen these libraries with DNA and RNA probes, and assemble and align extended regions of the chromosome of the bacterium Rhodobacter capsulatus. They use these methods to examine the process by which the bacterial genome undergoes evolutionary change. The approaches used in this course are widely used in eukaryotic genetics, such as in the identification of human disease genes and in the Human Genome Project. M. Fonstein. Spring. L.

216. Experimental Molecular Genetics. PQ: BioSci 200 or equivalent, or consent of instructor. This course satisfies two of the lab requirements (cell biology and genetics) for the biological sciences concentration. This course is designed to introduce students to the practice of research in molecular biology and genetics. An actual research topic is chosen with an attempt to obtain original, perhaps publishable, research results. Students are encouraged to make original contributions, including--in addition to executing experiments--experimental design, library searches, computerized sequence analysis, and written descriptions. Students cooperate in carrying out different aspects of the project. Previous topics included the isolation of new reporter genes and the development of new cloning vectors. M. Casadaban. Autumn. L.

217. General Genetics. PQ: BioSci 200 or equivalent, or consent of instructor. This is an advanced course in genetics designed to bridge the gap between BioSci 197 and graduate-level genetics that involves the application of molecular biology to some of the problems of classical genetics: the nature of the gene, recombination, and mutation. The role of mutation and recombination in immunology and carcinogenesis is discussed. The implications of our increasing knowledge of the base sequence of the human genome is considered in some detail. B. Strauss. Autumn.

218. Plant Genetics. PQ: Biological Sciences Sequence 1 or equivalent. This course assumes some knowledge of basic genetic principles and covers a number of topics of special interest that can be well illustrated with examples from plants. These topics include transmission genetics, genetic variation, allozymes, molecular genetic variation, genome organization and gene numbers, genetics of race and species differences, cytogenetics, polyploidy, translocation systems, asexual reproduction, mutation, genetic load, somaclonal variation, developmental genetics, heterosis, sex determination, self-incompatibility, cytoplasmic inheritance (chloroplast and mitochondial genomes), male sterility, transposable elements, disease and pest resistance, and genetic engineering. D. Charlesworth. Winter. Not offered 1995-96; will be offered 1996-97.

219. Human Genetics. PQ: Biological Sciences Sequence 1 or equivalent. Introduction to the methods of analysis of human genetic processes and the human genome, including molecular, pedigree, and population methods. These are applied to topics of social, medical, anthropological, and evolutionary interest. J. Spofford. Winter.

221. Human Developmental Biology. PQ: Biological Sciences Sequence 1 or equivalent and some knowledge of physics, chemistry, and organismal biology. This course fulfills the developmental biology requirement for the biological sciences concentration. This course examines the physiologic, cellular, and biochemical functions of a series of organs and systems in their transition from fetal to newborn life in the human and the implications of these changes for successful adaptation to independent life. Examples of failures of adaptation and disease states are presented and discussed. The organs and systems covered are brain, lung, heart, liver, immune system, blood-forming system, intestine, endocrine organs, and kidney. L. Gartner, Staff. Winter.

226. Animal Developmental Biology. PQ: Biological Sciences Sequence 1. This course fulfills the developmental biology requirement for the biological sciences concentration. This course studies developmental processes. Underlying mechanisms are illuminated through discussion of key experiments. Emphasis is on differentiation at different levels of development. Examples of developmental programs come from both invertebrate and vertebrate embryology. Subjects include pattern formation in the embryo, morphogenesis, cell and tissue interactions, and the control of gene expression in development. E. Ferguson, Staff. Spring. L.

227. Animal Developmental Biology. PQ: Biological Sciences Sequence 1. This course fulfills the developmental biology requirement for the biological sciences concentration. This course is identical to BioSci 226 except that it does not have a lab. E. Ferguson, Staff. Spring.

229. Plant Development and Molecular Genetics (=DevBio 329, Ec-Ev 329, EvBiol 329, MG/CB 361). PQ: Common Core biology. This course fulfills the developmental biology requirement for the biological sciences concentration. This course describes the growth, differentiation, and development of plants at the organismal, cellular, and molecular levels. Emphasis is placed on the regulatory function of plant hormones, particularly in response to environmental stimuli and in control of gene expression. Recent advances using molecular genetic approaches in Arabidopsis and maize will be a central feature of the course. M. Ruddat, B. Keith. Spring.

232. Mammalian Biology. PQ: Common Core biology. This course fulfills the organismal biology requirement for the biological sciences concentration. This course covers the structure and function of major organ systems of the typical mammal, with dissection, histological material, and lectures correlating function with gross and microscopic structure. Some focus on the organ systems of man. L. Straus, F. Straus. Autumn. L.

234. Chordate Biology. PQ: Common Core biology. This course fulfills the organismal biology requirement for the biological sciences concentration. This is a general consideration of the structure, evolution, phylogeny, and life history of vertebrates, with emphasis on comparative morphology and structural and functional evolution. J. Hopson. Autumn. L.

236. Evolution and Paleobiology. PQ: Common Core biology. This course fulfills the organismal biology requirement for the biological sciences concentration. Contemporary themes in evolution and paleobiology are presented in an interactive class format. Topics include the evolution of evolutionary thinking, recent models showing how evolution works, the great extinction controversy (climate, volcanoes, and asteroids), the nuts and bolts of reconstructing an evolutionary tree, and whether or not ontogeny recapitulates phylogeny. The lab provides basic background in paleontology and geology in preparation for an optional field trip during spring break to the Badlands of Big Bend National Park in southern Texas. Paleontologic topics include major events in the fossil record and dinosaur anatomy. Geologic topics include mineral and rock identification, stratigraphic principles, and the geology of Big Bend National Park. P. Sereno. Winter. L.

238. Introduction to Invertebrate Biology. PQ: Common Core biology or consent of instructor. This course fulfills the organismal biology requirement for the biological sciences concentration. This is a survey of the diversity, structure, and evolution of the invertebrate phyla, with emphasis on the major living and fossil invertebrate groups. Structure-function relationships and the influence of body plans on the evolutionary history of the invertebrate phyla are stressed. M. LaBarbera. Spring. L.

239. Prokaryotic Biology. PQ: 200-level course in cell biology or genetics. This course is an introduction to microbial structure and function, with an emphasis both on unique features and on those shared with eukaryotic forms. R. Haselkorn. Spring.

240. Biology and Evolution of Plants. PQ: Common Core biology. This course fulfills the organismal biology requirement for the biological sciences concentration. The lectures address the diversity in morphology, anatomy, reproduction, and evolutionary trends, beginning with cyanobacteria and progressing to flowering plants. The unifying aspects of cell structure and function are emphasized along with the basic physiological and molecular mechanisms in plants. The lab is correlated with the lectures to examine representatives of the major taxonomic plant groups and basic physiological techniques. M. Ruddat. Winter. L.

241. Biology and Evolution of Plants. PQ: Common Core biology. This course fulfills the organismal biology requirement for the biological sciences concentration. This course is identical to BioSci 240 except that it does not have a lab. M. Ruddat. Winter.

242. Physiology. PQ: Biological Sciences Sequence 1 or consent of instructor. This course fulfills the organismal biology requirement for the biological sciences concentration. This course is an intensive introduction to the mechanisms that operate in living organisms at all levels, ranging from the subcellular to the whole organism, to support organismal function. The course considers (1) molecular aspects of physiology (e.g., membrane function, channels, and receptors); (2) the neural and hormonal mechanisms that coordinate function; (3) muscle function and its regulation; and (4) the regulation of respiratory gas transport, temperature, water, and ions. A weekly lab illustrates principles introduced in the lecture. Staff. Winter. L.

244. Introduction to Neurobiology (=Biopsy 207). PQ: Common Core biology. This course fulfills the organismal biology requirement for the biological sciences concentration. An introduction to the diverse levels of analysis of the nervous system. Topics covered include structure of the nervous system, basic cellular neurobiology including membrane properties, synaptic transmission and transmitter/receptor systems, sensory transduction, nerve and muscle, central pattern generators, sensory systems, developmental neurobiology, and neuroethology. Comparative aspects of neurobiology are stressed, and both vertebrate and invertebrate examples are developed. D. Margoliash, M. Dickinson. Autumn. L.

246. Introductory Paleontology (=GeoSci 223). PQ: GeoSci 131-132, or Biological Sciences Sequence 1, or consent of instructor. This course fulfills the organismal biology requirement for the biological sciences concentration. The focus of the course is on the nature of the fossil record, the information it provides on patterns and processes of evolution through geologic time, and how it can be used to solve geological and biological problems. Lectures cover the principles of paleontology (including fossilization, classification, morphologic analysis and interpretation, biostratigraphy, paleoecology, and macroevolution); labs are systematic, introducing major groups of fossil invertebrates. J. Sepkoski. Winter. L.

248. Animal Behavior. PQ: Common Core biology. This course fulfills the ecology, populations, and behavior requirement for the biological sciences concentration. This is an introduction to the mechanisms and evolution of behavior in nonhuman species. The course is concerned with the mechanistic basis of behavior and the ecology and evolution of behavior at the individual and group level. Topics include behavioral genetics, developmental pathways, communication, foraging behavior, kin selection, mating systems, sexual selection, the ecological and social contexts of behavioral variation, and field and lab techniques. The lab focuses on research design and observational methods in behavioral biology and meets at Brookfield Zoo; transportation will be provided. J. Altmann, S. Pruett-Jones. Spring. L.

249. Animal Behavior (=Psych 214). PQ: Common Core biology. This course fulfills the ecology, populations, and behavior requirement for the biological sciences concentration. This course is identical to BioSci 248 except that it does not have a lab. S. Pruett-Jones, J. Altmann. Spring.

250. Evolutionary Ecology. PQ: Common Core biology. This course fulfills the ecology, populations, and behavior requirement for the biological sciences concentration. An evolutionary approach to the study of ecological interactions. Topics include plant-animal interactions, life history evolution, host-parasite and host-mutualist interactions, competition, and predation. Weekly labs stress experimental methods and exploration of current literature. E. Simms, M. Leibold. Winter. L.

251. Ecological Applications to Conservation Biology. PQ: BioSci 193 or 195 or consent of instructor. This course fulfills the ecology, population, and behavior requirement for the biological sciences concentration. We focus on the contribution of ecological theory to understanding current issues in conservation biology. The course emphasizes quantitative methods and their use for applied problems in ecology, such as the design of natural reserves, the risk of extinction, the impact of harvesting, and the dynamics of species invasions. Course material is drawn mostly from the current primary literature. In addition to lectures, students participate in weekly discussion groups and computer modeling lab exercises. J. Bergelson, C. Pfister, T. Nagylaki. Winter.

252. Field Ecology. PQ: Biological Sciences Sequence 1 and consent of instructor. This course fulfills the ecology, populations, and behavior requirement for the biological sciences concentration. This course is an introduction to habitats and biomes in North America and the methods of organizing and carrying out field research projects in ecology and behavior, focusing on questions of evolutionary significance. The course consists of a two-week field trip to the southwestern United States during the winter/spring quarter break. Work during the field trip consists of informal lectures and discussions, individual study, and group research projects. During the spring quarter there are lectures on the ecology of the areas visited and on techniques and methods of field research. This course is designed for students with a serious commitment to pursuing graduate research. S. Pruett-Jones. Spring. L. Not offered 1995-96; will be offered 1996-97.

254. Systematic Biology (=EvBiol 354). PQ: Common Core biology and knowledge of algebra. This course fulfills the ecology, populations, and behavior requirement for the biological sciences concentration. Systematic biology encompasses such activities as discovering and classifying biological diversity, estimating the phylogenetic relationships among species or larger lineages, and estimating evolutionary processes. From the standpoint of the three schools of systematic biology (evolutionary, phenetic, and phylogenetic), the course explores carefully the concepts of homology, species, and higher taxa. We consider the central role of systematic biology in the biological sciences and use systematic hypotheses to test theories about evolutionary or biological processes. B. Chernoff. Autumn. L.

255. Biogeography (=EvBiol 455, Geog 255/355). PQ: Common Core biology or consent of instructor. This course fulfills the ecology, populations, and behavior requirement for the biological sciences concentration. This course examines factors governing the distribution and abundance of animals and plants. Topics include patterns and processes in historical biogeography, island biogeography, geographical ecology, areography, and conservation biology (the design and effectiveness of nature reserves). Staff. Winter.

256. Fundamentals of Molecular Evolution. PQ: Biological Sciences Sequence 1 and calculus; or consent of instructor. The comparative analysis of DNA sequence variation has become an important tool in molecular biology, genetics, and evolutionary biology. This course covers major theories that form the foundation for understanding evolutionary forces governing molecular variation and divergence and genome organization. It explores the evolutionary assembly of genes, the origin of novel gene function, the population genetics of repetitive DNA variation, and the evolution of multigene families. The course also provides practical information on accessing genome databases, searching for homologous sequences, aligning DNA and protein sequences, calculating sequence divergence, producing sequence phylogenies, and estimating evolutionary parameters. M. Kreitman, T. Nagylaki. Spring.

259. Molecular Biocomputing. PQ: Biological Sciences Sequence 1 or equivalent; some knowledge of computing. This course introduces students to the various facets of using computers to augment research in molecular biology. Topics discussed may include experiment planning, expert systems, DNA/RNA/protein sequence analysis, and database access and usage. Issues such as the human genome project, genetic counseling, and DNA forensic science are also considered. This is a hands-on course with a significant lab component. J. Kruper. Spring. L.

260. Mammal Evolution (=EvBiol 311). PQ: Common Core biology or consent of instructor. This course fulfills the organismal biology requirement for biological sciences concentration. This course is an introduction to the major features of mammalian evolution. It surveys major groups of mammals, including both living and fossil taxa. We focus on phylogeny, morphology, biogeography, and patterns of diversification and extinction, using illustrations from the Field Museum's world-class collections of fossil and living mammals. Transportation to and from the museum is arranged as needed. J. Flynn. Autumn. L.

297. Readings in Biology. PQ: Consent of faculty sponsor. Students are required to submit the College Reading and Research Course Form. This is a tutorial offering individually designed readings. Students may take only one unit of BioSci 297 per quarter and must register by the end of the second week. This course is graded P/N and may not be used to satisfy an area requirement in advanced-level biology or as a substitute for one of the quarter courses in a Common Core biological sciences sequence. Staff. Summer, Autumn, Winter, Spring.

298. Undergraduate Research Seminar. PQ: BioSci 299. This seminar course is required of all graduating students who have taken BioSci 299 during the academic year and who plan to graduate with honors in the biological sciences and is open without credit to students who have taken BioSci 199. The poster session is held in early May. This course is graded P/F and may not be used to satisfy an area requirement in advanced-level biology or as a substitute for one of the quarter courses in a Common Core biology sequence. Staff. Spring.

299. Advanced Research in the Biological Sciences. PQ: Consent of faculty sponsor and the Undergraduate Research and Honors Committee. Students are required to submit the College Reading and Research Course Form. Advanced, individually guided research for undergraduate biology concentrators. Students submit a written report covering their research activities to the chairman of the Undergraduate Research and Honors Committee. As registrants in BioSci 299, they are required to participate in a biweekly senior honors forum beginning in autumn quarter and to register for BioSci 298 (Undergraduate Research and Seminar) in the spring quarter of their senior year. Both courses are graded P/F and may not be used to satisfy an area requirement in advanced-level biology or as a substitute for one of the quarter courses in a Common Core biology sequence. Staff. Summer, Autumn, Winter, Spring. L.

Specialized Courses

261. Viruses of Eukaryotes. PQ: Consent of instructor. This course is concerned with various aspects of the molecular biology of viruses of animal cells, including viruses that afflict man. Special emphasis is given to recent developments in the field related to viral nucleic acid replication, controls of viral gene expression, use of viruses as cloning vectors to amplify specific cellular genes, and the contribution of virus research to our understanding of mechanisms underlying eukaryotic gene expression. The course attempts to develop experimental thinking and knowledge of experimental approaches currently in use in related fields in molecular biology and cell biology. B. Roizman. Spring.

262. Electron Microscopy and Image Processing in Structural Biology (=MG/CB 310). PQ: One year of calculus. Electron microscopy is an important tool for studying the structure of biological macromolecular assemblies. Much information inherent in an electron micrograph is accessible only after the micrograph is subjected to a computer analysis of its periodic features. This course deals with the principles involved in processing electron microscope images, including the underlying analytical methods and their computer implementation. Students have an opportunity to use computers to analyze various periodic structures and to become acquainted with the various graphics systems used in image analysis. R. Josephs. Spring.

264. Endocrinology. PQ: Common Core biology or consent of instructor. A general introduction to the study of hormones and the homeostatic interaction of organ systems mediated by hormones. J. Jarabak, T. Jones. Winter.

265. The Biology of Toxoplasma (=Immun 365). This course undertakes a study of Toxoplasma gondii and toxoplasmosis: a model system to study the cellular and molecular biology, biochemistry, and genetics of an obligate intracellular protozoan parasite; the immune responses it elicits; and the pathogenesis of the diseases it causes. This information is also applied to consideration of public health measures for prevention of infection, for vaccines, and for development of new antimicrobial therapy. General principles applicable to the study of other microorganisms are emphasized. This course is suitable for undergraduates with a good background in biology and molecular genetics, as well as for graduate and medical students and postdoctoral fellows in immunology and infectious diseases. R. McLeod. Spring.

268. Neuropsychopharmacology (=Neurbi 327, PhaPhy 327, Psych 327). PQ: BioSci 200 or BchMB 301, or consent of instructor. This course entails a study of the effects of pharmacological agents on behavior with emphasis on physiological and biochemical mechanisms. L. Seiden, H. De Wit, P. Vezina. Autumn. L.

270. The Conquest of Pain. PQ: Organic chemistry or biochemistry; neurobiology or physiology recommended. This course examines the biology of pain and the mechanisms by which anesthetics alter the perception of pain. The approach is to examine the anatomy of pain pathways both centrally and peripherally, and to define electrophysiological, biophysical, and biochemical explanations underlying the action of general and local anesthetics. The role of opiates and enkephalins is discussed in detail. Central theories of anesthesia, including the relevance of sleep proteins, are also examined. Additionally, mechanistic discussions of acupuncture and cutaneous nerve stimulation are included. J. Moss. Spring.

272. Diet and Behavior. PQ: Common Core biology sequence. Students must attend the first class to confirm enrollment. No exceptions. Behavioral factors influence food selection and eating patterns, thus playing a key role in the process of facilitating dietary change. Disorders of eating may frequently be associated with changes in mood. This course utilizes weekly didactic sessions and the ambulatory clinical nutrition setting to examine topics in diet and behavior. In addition, case studies are utilized extensively to help explore pertinent issues. Since the subject matter is complex and extensive, students are required to review a large body of literature. M. H. Maskay, Staff. Autumn.

273. Evolutionary Processes (=CFS 348, Ec-Ev 310, EvBiol 310). PQ: Consent of instructor. Examination of evolutionary aspects of ecology, genetics, biochemistry, paleontology, development, philosophy, and related subjects through readings, written essays, and discussions. L. Van Valen. Winter.

275. Introduction to Psychiatry. PQ: Common Core biology. This course surveys fundamental aspects of the treatment, assessment, diagnosis, etiology, and prognosis of common psychiatric disorders. Emphasis is placed on the integration of epidemiologic, biological, psychological, and social perspectives in understanding and intervening in these disorders. Topics include principles of assessment and diagnosis, reviews of major psychiatric syndromes, and treatment modalities. This course also includes the historical and cultural context in which the contemporary practice of psychiatry takes place. Emphasis is placed on the history of concepts and institutions. S. Gilman. Winter.

276. Muscle Physiology and Function (=Med 510). PQ: Knowledge of physical concepts; elementary knowledge of chemistry and biology helpful; third- or fourth-year standing. This course considers muscle architecture, muscle structure, mechanisms of contraction, mechanochemical energy transduction, mechanisms of muscle activation, cardiac mechanics, animal locomotion, and optimization of locomotive parameters for athletics. L. Ford. Autumn.

278. Molecular Physiology. PQ: BioSci 242 or 243, or consent of instructor. This course examines insights yielded by the utilization of molecular techniques in physiological investigations. R. Zak, Staff. Not offered 1995-96; will be offered 1996-97.

281. Searching Bibliographic Databases (=HiPSS 221). Computer databases of indexes, abstracts, citations, and text provide indispensable routes of intellectual access to the world of knowledge. We focus on search strategy development, the organization and structure of databases and index languages, and techniques for evaluating search results. The interplay of human judgment with computer procedures is analyzed throughout, particularly in comparing subject, text, and citation searching. Using campus computers and the Internet, students search local and remote databases for references in journal articles related to a variety of assigned problems in the medical, biological, and environmental sciences and individually chosen problems that may be in any discipline. D. Swanson. Autumn, Spring. Not offered 1995-96; will be offered 1996-97.

Courses on the Ethical and Societal Implications of the Biological Sciences

282. Laboratory Fundamentals in Clinical Research. PQ: Common Core biology. This unique, new course has been designed to provide students in different stages of education and career development with the background necessary to plan, manage, and communicate within the world of clinical research. The course introduces students to the basics of the clinical laboratory and includes the tools, techniques, and skills required to assist in clinical human research protocols. Topics emphasized are lab safety, instrumentation, quality control and assurance, immunological assays, DNA preparation, and the ethics and logistics of research protocol development. The course consists of hands-on lab participation, informal lectures, and open discussion. S. Patel. Winter.

283. Disease (=Pathol 313). Intended for both science and nonscience concentrators, this course examines disease as a contextual and historical entity in the writings of several novelists and philosophers. S. Meredith. Spring.

284. The Quest for Human Nature. PQ: Common Core biology or consent of instructor. Do we have a nature? If so, what is it? How do we know? In this course, we join the enduring quest for human nature by exploring works of two of its seminal students, Aristotle and Darwin. What is the relationship between our biology and the ethical and social spheres of human life? How does our method of inquiry into human nature impact upon our results? In what ways might we manipulate our nature, and toward what ends might we change it? Our readings are drawn from select works by each author. R. Gunderman. Autumn. Not offered 1995-96; will be offered 1996-97.

285. Biological Approach to the Problem of Knowledge: Kant's Theory of Cognition (=HiPSS 285). PQ: Common Core biology. Term paper required for letter grade. This course is about human cognition, how the mind works in furnishing the kind of experience and knowledge which human beings have. It is about sensation and thought, consciousness and awareness of self, and, most importantly, about the reflections of Immanuel Kant on these matters. During the quarter, we acquire, stepwise, a grasp of Kantian theory by a close reading of the text of the Critique of Pure Reason, together with a few landmarks from the related scientific literature subsequent to Kant's time. The class sessions consist of detailed discussion of the texts, with students participating at a maximum level consistent with progress at a reasonable pace. S. Schulman. Autumn, Spring.

286. Foundation of Gender and Gender Differences (=NCD 228, Psych 219). This course examines issues fundamental to an understanding of sex, sex differences, and gender differences. The course begins by considering the significance of the evolution of sexually reproducing species and by studying mechanisms of normal and abnormal early embryonic sexual development. The relative influence of innate (or "primarily biological") and learned (or "primarily environmental") factors is discussed. The different reproductive roles of males and females in various species are examined to relate sexual dimorphism, mating strategies, and life cycle differences. This is a seminar course involving faculty from different departments. M. Moscona, M. McClintock. Spring.

290. Context of Medicine I (=Peds 342, PubPol 481, SSA 497). PQ: Consent of instructor. NOTE: This course ends in the spring quarter. Students receive a grade of Incomplete for the winter quarter and a course grade at the end of the spring quarter. The course considers the multiple influences on medicine and health care in the United States. It examines the roles of patients, physicians, and other health care professionals and institutions from historical, sociologic, economic, and interactional perspectives. Students participate in a variety of field experiences in hospitals and other settings to sharpen their skills as observers and data gatherers, and to have direct insight into the context of medicine. A. Kohrman, C. Kohrman. Winter.

291. Medicine and Ethics (=NCD 218). Clinical medicine and biomedical science pose fascinating questions. These involve permanent human concerns, such as the meaning of life, suffering, the relationship between healer and patient, and the pursuit of health, as well as issues on the cutting edge of contemporary science and society, such as new genetic and reproductive technologies, AIDS, and the organization and funding of health care. In the interplay between such questions lies not only enhanced understanding of pressing personal and public policy problems but also enduring insights into humanity. R. Gunderman. Autumn.

292. Medical Odysseys. PQ: Consent of instructor. Physicians and patients have new moral responsibilities due to changes in medical technology, economics, and public policy. Both physicians and patients must frame responses to the moral dilemmas of modern medicine: truth; conflict of interest; disparities in knowledge and power; allocation of scarce resources; and the meaning of life, disease, and death. This course studies works that present these and other dilemmas through the immediacy of lived personal experiences, as documented in books of medical autobiography, essays, and poems. J. Lantos, A. Goldblatt. Winter. Not offered 1995-96; will be offered 1996-97.

294. Professional Ethics in Science (=HiPSS 297). PQ: Common Core biology or consent of instructor. This course is designed primarily for students pursuing a professional career in science rather than medicine. It explores the ethical context of biology by analyzing the values, goals, purposes, moral claims, and aspirations of the professional scientific activity. It does not present scientific material but draws heavily on the works of major figures who developed systems of ethics and of scientists who have articulated the ethos of science as actually practiced. These theoretical systems are applied to a few paradigmatic case studies in which societal values seem to be in conflict with the scientific ethic. The course concludes with a presentation of a major theory in which human values are given a biological foundation. R. Holmes. Spring.

296. The New Genetics and Health Care. PQ: Consent of instructor. This course addresses issues raised by research in genetics and examines the implications of alternative solutions. Topics to be considered are the goals, methods, and achievements of the Human Genome Project; examples of genetic conditions about which information is being or will be generated; implications of genetics for concepts of disease and health; cultural, ethnic, gender, and socioeconomic differences in genetic research and its applications; ethical and legal issues related to genetic tests and screening; genetic counseling and medical models of the clinician-patient interaction; ethical and policy dimensions of genetic disabilities; and scientific, clinical, ethical, and legal aspects of gene theory. M. Mahowald, M. Verp. Autumn.

Graduate-Level Courses

Many graduate-level courses in the Division of Biological Sciences are open to qualified College students. Students should consult their advisers, the BSCD office, or the various departments and committees in the division to identify appropriate courses.

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