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Go to: Program Requirements
Cell Biology, Molecular Biology, and Genetics. Cells are fundamental units
of biological organization. The properties of cells are determined by the genes
they contain and express. The recent development of methods to isolate and
clone genes, to mutate these genes in vitro, and to transfer these genes into
other cells or organisms, is revolutionizing our understanding of biological
processes. Courses in cell biology, molecular biology, and genetics provide a
key to understanding the profound ways that these biological discoveries will
affect our lives.
Organismal Biology. Multicellular organisms comprise many specialized types
of cells. The survival and reproduction of an organism necessitate the
coordination and integration of the functions of its component cells. The
functions of cells, and therefore of organisms, are dependent upon a continual
exchange of matter and energy between the organism and its environment. Courses
in organismal biology address these problems and their solutions in animals and
plants from a variety of perspectives.
Population Biology, Evolution, Ecology, and Behavior. Evolution is perhaps
the most important unifying concept in the biological sciences. Natural
populations are characterized by genetic and phenotypic diversity. This
diversity forms the basis for evolution by natural selection. Selection
involves interactions between organisms and their environment. Courses in
population biology and evolution consider the evidence for and the mechanisms
of evolution and the ways in which evolutionary and ecological processes affect
populations and species.
Each Common Core course includes laboratory exercises. In these laboratories,
students gain first-hand experience in carrying out experiments or making
observations and in analyzing and interpreting the biological data they obtain
from these experiments and observations. The laboratories are an integral part
of the courses; to receive a grade in a Common Core course, students must
complete the laboratory exercises associated with that course even if the
course is taken as an elective. Some courses include laboratories that require
the use of animals. Students who have concerns about using animals in
laboratories are urged to discuss these concerns with the instructors before
registering for those courses.
Common Core courses vary with respect to specific subject matter and format.
This variety offers students the opportunity to choose a Common Core sequence
suited to their experience, interests, and educational goals. To choose the
most appropriate Common Core sequence, students should consult the faculty, the
College advisers, fellow students, and the Student Course Evaluations
book.
Students who concentrate in the biological sciences are required to complete
Sequence 1. Students may use three of these courses to fulfill their Common
Core requirements, or they may take another Common Core sequence and use
Biological Sciences Sequence 1 to fulfill part of their concentration
requirement. Taking Sequence 1 as the Common Core enables students to take more
advanced biology courses without using electives. On the other hand, taking
another Common Core sequence provides students the opportunity to explore
diverse areas of biology early in their program and may help students who are
uncertain about concentrating in biology to make a more informed decision. In
the past, approximately 60 percent of biological sciences concentrators have
taken Sequence 1 as their Common Core while 40 percent have taken other Common
Core sequences.
Common Core courses are designed to fulfill the general education requirement
in the biological sciences and to prepare students for entering the biological
sciences concentration. Students who have taken or are currently taking
Biological Sciences Sequence 1 may not enroll in other Common Core courses
without the consent of the senior adviser.
Placement. Students who score 4 or 5 on the AP test in biology, or who
perform well on the College's optional placement test in the biological
sciences, will be excused from the Common Core requirement in the biological
sciences. Students who place out of the biology Common Core and who
subsequently concentrate in the biological sciences are required either to
complete Biological Sciences Sequence 1 as part of their concentration
requirement or demonstrate mastery of the material in Biological Sciences
Sequence 1 by examination. Students who wish to take the placement exam for
Biological Sciences Sequence 1 should consult the senior adviser.
Accreditation. Students who place out of the Common Core biology sequence
may elect to take an accreditation examination in one or more of the five areas
of required advanced biology (biochemistry, cell biology, developmental
biology, organismal biology, and evolution) during the first year of
registration. Credit is granted upon satisfactory completion of an oral
examination before three members of the faculty designated for each area by the
BSCD master. No laboratory requirements can be met by accreditation
examinations except by special petition with accompanying documentation.
Accreditation for advanced biology courses may also be granted to students at
any time after completion of the 100-level Common Core biology sequence and
upon satisfactory completion of the same oral examination. No oral examination
for accreditation may be taken more than once.
Students are urged to consult the BSCD senior adviser immediately if they feel
that they can qualify for accreditation.
The program for a B.A. in the biological sciences consists of forty-two
credits, including the following:
1. The Common Core. To prepare themselves for more advanced work in
biology, prospective biological sciences concentrators should take Chemistry
111-112-113 or higher to satisfy the Common Core requirement in the physical
sciences, and Mathematics 131-132 or higher to fulfill the mathematics
requirement. Students who concentrate in the biological sciences may satisfy
their Common Core requirement in the biological sciences with any Common Core
biological sciences sequence; however, they must complete Biological Sciences
Sequence 1 either as their Common Core requirement or as part of their
concentration program.
2. Other Science Courses. Biological sciences concentrators must
complete a three-quarter course in physics (usually Physics 121-122-123 or
higher), Chemistry 217-218 or Chemistry 220-221, and Statistics 220.
3. Advanced-Level Biology Courses. Each student must take the following
advanced biology courses:
a. Biological Sciences Sequence 1 (four courses). If students take three
quarters of Sequence 1 to fulfill their Common Core requirement, they then have
the opportunity to take three advanced courses in addition to the fourth
quarter of Sequence 1. One of these courses must be in ecology, population
biology, or behavior (Biological Sciences 248-255). The others may be two
200-level courses in science (biology or chemistry) or one course in science
and one in mathematics. A third quarter of calculus is strongly recommended.
b. Biochemistry (Biological Sciences 200). Students are encouraged to take
Biological Sciences 200 in the spring quarter of the second year or the autumn
quarter of the third, if possible, so that they can take cell biology
(Biological Sciences 207) in their third year. Students may also satisfy this
requirement by completing both Biochemistry and Molecular Biology 301 and 302;
students who do this must take 302 as an elective.
c. Cell biology (Biological Sciences 207) and developmental biology (one of
Biological Sciences 221-229). Students who took Biological Sciences 196-197 in
1994-95 may take two elective 200-level biological sciences courses in lieu of
these requirements.
d. Organismal biology (one of Biological Sciences 232-246, or 260).
4. Laboratories. Students must take three 200-level biological sciences
courses with a laboratory. Courses used to satisfy the laboratory
requirement need not be the same courses used to satisfy other concentration
requirements, but they can be. The three laboratory courses must be in
different areas, as follows:
a. Biochemistry (Biological Sciences 200 or 259).
b. Cell biology (Biological Sciences 208, 215, or 216).
c. Developmental biology (Biological Sciences 226).
d. Genetics (Biological Sciences 215, 216, or 220).
e. Neurobiology (Biological Sciences 212, 244, or 268).
f. Organismal biology (Biological Sciences 232, 234, 236, 238, 240, 242, 246,
or 260).
g. Ecology, population biology, and behavior (Biological Sciences 248, 250,
252, or 254).
h. Biocomputing (Biological Sciences 267).
Biological Sciences 215 and 216 each satisfy two of the laboratory
requirements (cell biology and genetics) for the biological sciences
concentration. In addition, graduate courses with laboratory can usually be
used to satisfy these requirements; consult with the senior adviser. Please
note, however, that Biological Sciences 199, 297, 298, and 299 cannot be used
to satisfy the laboratory requirement.
5. Free Electives. Students have up to seven free electives. They may
use these electives to engage in honors research (Biological Sciences 199, 298,
and 299), to take directed reading courses in biology (Biological Sciences
297), or to take additional courses in biology or other subjects of their
choice.
6. Grading. Students must receive quality (letter) grades in all
fourteen courses in the concentration.
Concentration
4 BioSci Sequence 1. If three courses in Sequence 1 are taken
as Common Core, they may be replaced by three 200-level
courses in science (biology or chemistry) or two courses in
science and one in mathematics. One of these courses must
be in ecology, population biology, or behavior (BioSci 248-255).
*Students who took Biological Sciences 196-197 in 1994-95 may take two elective
200-level biological sciences courses in lieu of these requirements.
Research Courses. Students are strongly encouraged to carry out individual
guided research in an area of their interest. Students who choose to carry out
research can receive academic credit for their work and have the opportunity to
graduate with special honors. Any faculty member in the Division of the
Biological Sciences can sponsor and supervise student research on an individual
tutorial basis. The activities of the student are arranged by mutual agreement
between the student and the sponsor. Students can find out about research
opportunities and potential faculty sponsors from the College Research
Opportunities Program (CROP) Directory, available in the Office of the Dean
of the College, from the BSCD Office, or from the chairman of the Undergraduate
Research and Honors Committee.
Students can receive academic credit for their research by enrolling in
research courses. Two such courses are available: Biological Sciences 199
(Introduction to Research) and Biological Sciences 299 (Advanced Research in
the Biological Sciences). Biological Sciences 199 is intended for students who
wish to have a research experience but are not conducting an approved honors
research project; it may be elected for up to three quarters. Biological
Sciences 299 is for advanced students, including but not limited to those who
wish to be candidates for special honors. Registration for either Biological
Sciences 199 or Biological Sciences 299 requires permission of the student's
research sponsor and the chairman of the Undergraduate Research and Honors
Committee. To register for Biological Sciences 299, students must submit a
written research proposal to the chairman of the Undergraduate Research and
Honors Committee. A registration form for these courses may be obtained from
your adviser. Students must complete this form before registering for
research courses. Both Biological Sciences 199 and Biological Sciences
299 are graded P/F; neither course can be used to fulfill concentration
requirements.
Students who take Biological Sciences 299 are required to participate in a
biweekly senior honors forum beginning in autumn quarter. In addition, students
who wish to receive special honors must participate in Biological Sciences 298
(Undergraduate Research Seminar) in their senior year. In the honors forum and
seminar, students report on and discuss their research. Students who take
Biological Sciences 199 are encouraged to attend the forum and seminar but will
normally not present their work. Biological Sciences 298 is offered in the
spring quarter; students who cannot take this course because they plan to
graduate in a different quarter should contact the chairman of the
Undergraduate Research and Honors Committee at least three months before their
expected graduation to arrange for an equivalent experience. Biological
Sciences 298 is also graded P/F and may not be used to fulfill a
concentration requirement.
Special Honors. Students who wish to receive their bachelor's degree with
special honors in the biological sciences must complete an individual research
project. Students should discuss their plans with the chairman of the
Undergraduate Research and Honors Committee as early as possible. Many students
begin their honors research projects during the summer between their junior and
senior years. Every effort is made to provide financial support for students
who wish to carry out summer research. Students should normally begin their
research projects no later than the autumn quarter of their senior year.
Students who are working toward honors must participate in Biological Sciences
299 and Biological Sciences 298 and must complete a research project under the
supervision of a faculty sponsor. This research project may consist of either
experimental work or fieldwork. In the spring quarter of the senior year,
students must prepare and submit a paper based on the results of their research
and must present and defend their work, normally at a poster session open to
the general public. Approval of the paper and successful defense qualify a
student to receive special honors at graduation, provided he or she also has
the grade point average necessary for special honors (an overall grade point
average of 3.0 or an average of 3.25 in the fourteen-course concentration
program.)
This program requires the completion of forty-eight credits of course work
rather than the forty-two required for the B.A. (Actually, fifty-one
credits--forty-two for the B.A. and nine for the M.S.--are required; however,
up to three upper-level course credits can be double-counted for both degrees.)
Thus, the program is difficult in terms of the course load and also requires
payment of graduate-level tuition during the senior year. In general, the
program is suitable only for students who enter with placement or advanced
placement credit or who complete some of their work during the summer.
A departmental oral examination on a thesis describing the candidate's original
research contribution is required for the degree. Students are expected to (1)
spend elective time and/or a summer residency in a laboratory working on a
research problem in biochemistry or molecular biology, and (2) write a formal
thesis describing their research and reviewing the field. The oral examination
based on that thesis must be taken during the last quarter of their
registration.
Students will not be admitted formally to the program until their third year.
Applications are made through the Department of Biochemistry and Molecular
Biology. Normally only students with grade point averages above 3.0 qualify,
although exceptions may be made under special circumstances.
Questions about the program may be addressed to the adviser in the Department
of Biochemistry and Molecular Biology (Marvin Makinen, CLSC 359, 702-1080).
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JEANNE ALTMANN, Professor, Department of Ecology & Evolution, Committees on
Biopsychology and Evolutionary Biology, and the College; Chairman, Committee on
Evolutionary Biology
STEVAN J. ARNOLD, Professor, Department of Ecology & Evolution, Committees
on Evolutionary Biology and Genetics, and the College
R. DEAN ASTUMIAN, Assistant Professor, Departments of Biochemistry &
Molecular Biology and Surgery and the College
JUDITH AUSTIN, Assistant Professor, Department of Molecular Genetics & Cell
Biology, Committee on Developmental Biology, and the College
BARBARA L. BERG, Lecturer in the College
JOY M. BERGELSON, Assistant Professor, Department of Ecology & Evolution
and the College
RUDIGER BIELER, Lecturer, Committee on Evolutionary Biology and the College
ANDREW BIEWENER, Associate Professor, Department of Organismal Biology &
Anatomy, Committee on Evolutionary Biology, and the College; Chairman,
Department of Organismal Biology & Anatomy
JEFFREY A. BLUESTONE, Professor, Ben May Institute, Department of Pathology,
Committee on Immunology, and the College; Chairman, Committee on Immunology
JAMES E. BOWMAN, M.D., Professor Emeritus, Departments of Pathology and
Medicine, Committees on African & African-American Studies and Genetics,
and the College
DONN BRANSTRATOR, Lecturer in the College
MALCOLM J. CASADABAN, Associate Professor, Departments of Biochemistry &
Molecular Biology and Molecular Genetics & Cell Biology, Committees on
Genetics and Virology, and the College
BRIAN CHARLESWORTH, George Wells Beadle Distinguished Service Professor,
Department of Ecology & Evolution, Committees on Evolutionary Biology and
Genetics, and the College
DEBORAH CHARLESWORTH, Professor, Department of Ecology & Evolution,
Committees on Evolutionary Biology and Genetics, and the College
BARRY CHERNOFF, Lecturer, Committee on Evolutionary Biology and the College
KWEN-SHENG CHIANG, Associate Professor, Department of Molecular Genetics &
Cell Biology, Committee on Genetics, and the College
LIANE COCHRAN-STAFIRA, Lecturer in the College
THOMAS COLTON, Senior Lecturer in the College
JERRY COYNE, Professor, Department of Ecology & Evolution, Committees on
Evolutionary Biology and Genetics, and the College
MARY CRANE, Lecturer in the College
DOUGLAS CRAWFORD, Assistant Professor, Department of Organismal Biology &
Anatomy, Committees on Evolutionary Biology and Genetics, and the College
MICHAEL DICKINSON, Assistant Professor, Department of Organismal Biology &
Anatomy, Committee on Neurobiology, and the College
RONALD DUBREUIL, Assistant Professor, Department of Pharmacological &
Physiological Sciences, Committee on Cell Physiology, and the College
WOLFGANG EPSTEIN, M.D., Professor, Departments of Biochemistry & Molecular
Biology and Molecular Genetics & Cell Biology, Committee on Genetics, and
the College
ROCHELLE EASTON ESPOSITO, Professor, Department of Molecular Genetics &
Cell Biology, and the College; Chairman, Committee on Genetics
JULIANA H. FEDER, Research Associate (Instructor), Department of Molecular
Genetics & Cell Biology and Lecturer in the College
MARTIN E. FEDER, Professor, Department of Organismal Biology & Anatomy,
Committee on Evolutionary Biology, and the College
EDWIN FERGUSON, Assistant Professor, Department of Molecular Genetics &
Cell Biology, Committees on Developmental Biology and Genetics, and the
College
EILEEN FIELDING, Lecturer in the College
FRANK W. FITCH, M.D., Albert D. Lasker Professor, Department of Pathology,
Committees on Developmental Biology and Immunology, and the College; Director,
Ben May Institute
JOHN FLYNN, Lecturer, Committee on Evolutionary Biology and the College
MICHAEL FONSTEIN, Research Associate (Assistant Professor), Department of
Molecular Genetics & Cell Biology; Lecturer in the College
LINCOLN E. FORD, M.D., Associate Professor, Department of Medicine and the
College
HARRY A. FOZZARD, M.D., Otho S. A. Sprague Distinguished Service Professor,
Departments of Medicine and Pharmacological & Physiological Sciences,
Committee on Cell Physiology, and the College; Chairman, Department of
Pharmacological & Physiological Sciences
HERBERT C. FRIEDMANN, Associate Professor, Department of Biochemistry &
Molecular Biology and the College
EDWARD D. GARBER, Professor Emeritus, Department of Ecology & Evolution,
Committee on Genetics, and the College
LAWRENCE M. GARTNER, M.D., Professor, Department of Pediatrics and the
College
GODFREY S. GETZ, M.D., Donald N. Pritzker Professor, Departments of
Biochemistry & Molecular Biology, Medicine, and Pathology, and the College;
Acting Dean, Division of the Biological Sciences and the Pritzker School of
Medicine; Chairman, Department of Pathology
SANDER L. GILMAN, Professor, Departments of Germanic Studies and Psychiatry,
and the College
ANN D. GOLDBLATT, Lecturer, Department of Medicine and the College
LARRY GOLDMAN, M.D., Associate Professor, Department of Psychiatry and the
College
EUGENE GOLDWASSER, Alice Hogge and Arthur A. Baer Professor Emeritus,
Department of Biochemistry & Molecular Biology, Committee on Developmental
Biology, and the College; Chairman, Department of Biochemistry & Molecular
Biology
DANIEL GOTTSCHLING, Associate Professor, Department of Molecular Genetics &
Cell Biology, Committee on Genetics, and the College
GEOFFREY GREENE, Professor, Ben May Institute, Department of Biochemistry &
Molecular Biology and the College
RICHARD GUNDERMAN, Lecturer in the College
DOROTHY A. HANCK, Assistant Professor, Department of Medicine, Committees on
Cell Physiology and Neurobiology, and the College
ROBERT HASELKORN, Fanny L. Pritzker Distinguished Service Professor,
Departments of Biochemistry & Molecular Biology, Chemistry, and Molecular
Genetics & Cell Biology, Committees on Developmental Biology, Genetics, and
Virology, and the College
ALFRED HELLER, M.D., Professor, Department of Pharmacological &
Physiological Sciences and the College
PHILIP C. HOFFMANN, Professor, Department of Pharmacological &
Physiological Sciences and the College
H. RODNEY HOLMES, Senior Lecturer in the College
JAMES A. HOPSON, Professor, Department of Organismal Biology & Anatomy,
Committee on Evolutionary Biology, and the College
LESLIE E. HORNIG, Lecturer in the College
LYNNE HOUCK, Research Associate (Associate Professor), Department of Ecology
& Evolution; Lecturer in the College
JOSEPH JARABAK, M.D., Associate Professor, Department of Medicine and the
College
THOMAS M. JONES, M.D., Professor, Department of Medicine and the College
ROBERT JOSEPHS, Professor, Department of Molecular Genetics & Cell Biology
and the College
BRIAN KEITH, Assistant Professor, Department of Molecular Genetics & Cell
Biology, Committees on Developmental Biology and Genetics, and the College
ARTHUR F. KOHRMAN, M.D., Professor, Department of Pediatrics and the College
CLAIRE KOHRMAN, Research Associate (Associate Professor), Center for Health
Administration Studies and Department of Pediatrics; Lecturer in the College
MARTIN KREITMAN, Associate Professor, Department of Ecology & Evolution,
Committees on Evolutionary Biology and Genetics, and the College
JOHN KRUPER, Director, Academic Computing, Division of the Biological Sciences;
Lecturer in the College
ROBERT KUSHNER, M.D., Assistant Professor, Department of Medicine and the
College
MICHAEL C. LABARBERA, Professor, Departments of the Geophysical Sciences and
Organismal Biology & Anatomy, Committee on Evolutionary Biology, and the
College
THOMAS G. LAMMERS, Lecturer in the College
GAYLE LAMPPA, Associate Professor, Department of Molecular Genetics & Cell
Biology, Committee on Genetics, and the College
JOHN LANTOS, M.D., Associate Professor, Departments of Medicine and Pediatrics
and the College
MATHEW LEIBOLD, Assistant Professor, Department of Ecology & Evolution,
Committee on Evolutionary Biology, and the College
SCOTT LIDGARD, Lecturer, Committee on Evolutionary Biology and the College
SUSAN LINDQUIST, Professor, Howard Hughes Medical Institute, Department of
Molecular Genetics & Cell Biology, Committees on Developmental Biology and
Genetics, and the College
PHILIP E. LLOYD, Associate Professor, Department of Pharmacological &
Physiological Sciences, Committees on Cell Physiology and Neurobiology, and the
College; Chairman, Committee on Neurobiology
R. ERIC LOMBARD, Associate Professor, Department of Organismal Biology &
Anatomy, Committee on Evolutionary Biology, and the College
ANTHONY P. MAHOWALD, Louis Block Professor, Department of Molecular Genetics
& Cell Biology, Committees on Developmental Biology, Genetics, and
Neurobiology, and the College; Chairman, Department of Molecular Genetics &
Cell Biology and Committee on Developmental Biology
MARY MAHOWALD, Professor, Department of Obstetrics & Gynecology and the
College
MARVIN W. MAKINEN, Professor, Department of Biochemistry & Molecular
Biology and the College
DANIEL MARGOLIASH, Associate Professor, Departments of Organismal Biology &
Anatomy and Psychology, Committees on Biopsychology and Neurobiology, and the
College
ALVIN MARKOVITZ, Professor, Department of Biochemistry & Molecular Biology,
Committee on Genetics, and the College
PALMA ANN MARONE, Lecturer in the College
TERENCE E. MARTIN, Professor, Department of Molecular Genetics & Cell
Biology, Committees on Genetics and Immunology, and the College
MARK MARTINDALE, Assistant Professor, Department of Organismal Biology &
Anatomy, Committees on Developmental Biology and Neurobiology, and the
College
MANISHA H. MASKAY, Assistant Professor, Department of Medicine and the
College
JANE MASTERSON, Lecturer in the College
STEPHANIE MAYER, Research Associate, Department of Ecology & Evolution;
Lecturer in the College
MARTHA K. MCCLINTOCK, Professor, Department of Psychology, Committees on
Evolutionary Biology and Neurobiology, and the College; Chairman, Committee on
Biopsychology
RIMA MCLEOD, M.D., Lecturer, Department of Medicine, Committee on Immunology,
and the College
STEPHEN C. MEREDITH, M.D., Associate Professor, Department of Pathology and the
College
LAURENS METS, Associate Professor, Department of Molecular Genetics & Cell
Biology, Committee on Genetics, and the College
JAMES MILLER, Associate Professor, Departments of Molecular Genetics & Cell
Biology and Pathology, Committees on Developmental Biology and Immunology, and
the College
HOWARD MOLTZ, Professor, Department of Psychology and the College
JOSEPHINE A. MORELLO, Professor, Departments of Medicine and Pathology and the
College
MALKA MOSCONA, Associate Professor in the College
JONATHAN MOSS, M.D., Professor, Department of Anesthesiology & Critical
Care and the College
GREGORY H. MUELLER, Lecturer, Committee on Evolutionary Biology and the
College
MARK MUSCH, Research Associate (Assistant Professor), Department of Medicine;
Lecturer in the College
JAMES NACHMAN, M.D., Associate Professor, Department of Pediatrics and the
College
THOMAS NAGYLAKI, Professor, Department of Ecology & Evolution, Committees
on Evolutionary Biology and Genetics, and the College
MATTHEW NITECKI, Curator, Committee on Evolutionary Biology; Lecturer in the
College
AVERTANO NORONHA, M.D., Associate Professor, Department of Neurology, Committee
on Immunology, and the College
CAROLE OBER, Associate Professor, Department of Obstetrics & Gynecology,
Committees on Evolutionary Biology and Genetics, and the College
SUJATA PATEL, Lecturer in the College
BRUCE PATTERSON, Lecturer, Committee on Evolutionary Biology and the College
ROBERT L. PERLMAN, M.D., Professor, Departments of Pediatrics and
Pharmacological & Physiological Sciences, Committees on Cell Physiology and
Neurobiology, and the College
CATHY A. PFISTER, Assistant Professor, Department of Ecology & Evolution
and the College
ROSAMOND V. POTTER, Senior Lecturer in the College
STEPHEN PRUETT-JONES, Associate Professor, Department of Ecology &
Evolution, Committee on Evolutionary Biology, and the College
JOSE QUINTANS, M.D., Professor, Department of Pathology and the College;
Master, Biological Sciences Collegiate Division; Associate Dean, Division of
Biological Sciences and the College
HYMAN ROCHMAN, M.D., Associate Professor, Department of Pathology and the
College
BERNARD ROIZMAN, Joseph Regenstein Distinguished Service Professor, Departments
of Biochemistry & Molecular Biology and Molecular Genetics & Cell
Biology, Committees on Genetics and Virology, and the College; Chairman,
Committee on Virology
MARSHA R. ROSNER, Professor, Ben May Institute, Department of Pharmacological
& Physiological Sciences, Committee on Cell Physiology, and the College;
Chairman, Committee on Cancer Biology
LUCIA ROTHMAN-DENES, Professor, Department of Molecular Genetics & Cell
Biology, Committee on Genetics, and the College
MANFRED D. E. RUDDAT, Associate Professor, Department of Ecology &
Evolution, Committee on Developmental Biology, and the College; Associate Dean
of the College; Senior Adviser, Biological Sciences Collegiate Division
DALE A. SCHOELLER, Associate Professor, Department of Medicine and the College;
Chairman, Committee on Human Nutrition & Nutritional Biology
SIDNEY SCHULMAN, M.D., Ellen C. Manning Professor Emeritus, Division of the
Biological Sciences and the College
GEBHARD F. B. SCHUMACHER, M.D., Professor Emeritus, Department of Obstetrics
& Gynecology and the College
PAUL T. SCHUMACKER, Professor, Department of Medicine and the College
LEWIS S. SEIDEN, Professor, Departments of Pharmacological & Physiological
Sciences and Psychiatry, Committee on Neurobiology, and the College
J. JOHN SEPKOSKI, JR., Professor, Departments of the Geophysical Sciences and
Organismal Biology & Anatomy, Committee on Evolutionary Biology, and the
College
PAUL SERENO, Associate Professor, Department of Organismal Biology &
Anatomy, Committee on Evolutionary Biology, and the College
JAMES SHAPIRO, Professor, Department of Biochemistry & Molecular Biology,
Committee on Genetics, and the College
PETRA SIERWALD, Lecturer in the College
ELLEN SIMMS, Assistant Professor, Department of Ecology & Evolution,
Committee on Evolutionary Biology, and the College
NEIL R. SMALHEISER, M.D., Assistant Professor, Department of Pediatrics,
Committees on Developmental Biology and Neurobiology, and the College
PATRICIA A. SOLTYS, Lecturer in the College
JANICE B. SPOFFORD, Associate Professor, Department of Ecology & Evolution,
Committees on Developmental Biology, Evolutionary Biology, and Genetics, and
the College
THEODORE L. STECK, M.D., Professor, Departments of Biochemistry & Molecular
Biology and Molecular Genetics & Cell Biology, Committee on Cell
Physiology, and the College
DONALD F. STEINER, M.D., A. N. Pritzker Professor, Howard Hughes Medical
Institute, Departments of Biochemistry & Molecular Biology and Medicine,
and the College
URSULA STORB, Professor, Departments of Molecular Genetics & Cell Biology
and Pathology, Committees on Genetics, Developmental Biology, and Immunology,
and the College
FRANCIS H. STRAUS II, M.D., Professor, Department of Pathology and the
College
LORNA P. STRAUS, Professor, Department of Organismal Biology & Anatomy and
the College
BERNARD S. STRAUSS, Professor, Department of Molecular Genetics & Cell
Biology, Committee on Genetics, and the College
PAUL STRIELEMAN, Lecturer in the College
DON R. SWANSON, Professor, Division of the Humanities and the College
KEVIN SWIER, Lecturer in the College
SARA SZUCHET, Professor, Department of Neurology, Committee on Neurobiology,
and the College
EDWIN W. TAYLOR, Louis Block Professor, Departments of Biochemistry &
Molecular Biology and Molecular Genetics & Cell Biology, Committee on Cell
Physiology, and the College
FRANK K. THORP, Associate Professor, Department of Pediatrics, Committee on
Human Nutrition & Nutritional Biology, and the College
RUSSELL H. TUTTLE, Professor, Department of Anthropology, Committee on
Evolutionary Biology, Morris Fishbein Center for the History of Science &
Medicine, and the College
PHILIP S. ULINSKI, Professor, Department of Organismal Biology & Anatomy,
Committee on Neurobiology, and the College
LEIGH M. VAN VALEN, Professor, Department of Ecology & Evolution,
Committees on Conceptual Foundations of Science, Evolutionary Biology, and
Genetics, and the College
MARION VERP, Associate Professor, Department of Obstetrics & Gynecology and
the College
MITCHEL VILLEREAL, Professor, Department of Pharmacological & Physiological
Sciences, Committee on Cell Physiology, and the College; Chairman, Committee on
Cell Physiology
MICHAEL J. WADE, Professor, Department of Ecology & Evolution, Committees
on Evolutionary Biology and Genetics, and the College; Chairman, Department of
Ecology & Evolution
NORMA E. WAGONER, Professor, Department of Organismal Biology & Anatomy and
the College; Dean of Students, Division of the Biological Sciences and the
Pritzker School of Medicine
MARK WESTNEAT, Lecturer, Department of Organismal Biology & Anatomy,
Committee on Evolutionary Biology, and the College
HUGH R. WILSON, Professor, Department of Ophthalmology & Visual Science,
Committee on Neurobiology, and the College
WILLIAM WIMSATT, Professor, Department of Philosophy, Committees on Conceptual
Foundations of Science, Evolutionary Biology, and General Studies in the
Humanities, Morris Fishbein Center for the History of Science & Medicine,
and the College
J. TIMOTHY WOOTTON, Assistant Professor, Department of Ecology & Evolution,
Committee on Evolutionary Biology, and the College
RADOVAN ZAK, M.D., Professor, Departments of Medicine, Organismal Biology &
Anatomy and Pharmacological & Physiological Sciences, Committee on Cell
Physiology, and the College
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Go to: Summary of Requirements
Go to: Faculty
Biological Sciences
Master: José Quintans, BSLC 104A, 702-7964
Senior Adviser: Manfred D. E. Ruddat, HM 254, 702-8623
Administrative Assistant: G. Karen Shrode, BSLC 104C, 702-7962
Staff Secretary: Cynthia Piwowar, BSLC 104B, 702-7963
Faculty Advisers: James Hopson, Organismal Biology & Anatomy, A 405B, 702-8099; Laurens Mets, Molecular Genetics & Cell Biology, EBC 210, 702-8917; Stephen Pruett-Jones, Ecology & Evolution, Z 47, 702-3115; Marvin Makinen, Biochemistry & Molecular Biology, CLSC 359, 702-1080
Undergraduate Research and Honors Chair: To be announced
Program of Study
Biology is the study of living things. The faculty of the College believes that
a sound knowledge of biology is essential for understanding many of the most
pressing problems of modern life and for intelligent involvement in their
eventual solution. The Biological Sciences Collegiate Division, therefore,
provides a variety of Common Core courses for all College students, prospective
biologists and nonbiologists alike. Although most of the course offerings
beyond the introductory year are designed to serve the needs of biology
concentrators, many of these courses are well suited to students in other
concentrations who wish to study some aspect of modern biology in greater
detail. Courses on the ethical and societal implications of the biological
sciences, for example, will be of interest to all students.
The Biological Sciences Common Core
The goal of the Common Core biological sciences courses is to give students an
appreciation of the problems or questions with which the biological sciences
are concerned, the modes of inquiry by which biological scientists address
these problems, and the ways in which biologists analyze and interpret the
results of this inquiry. At the same time, the Common Core curriculum gives
students a sufficient understanding of the current "biological revolution,"
enabling them to participate fully as informed citizens in a society that is
increasingly affected by the fruits of this revolution. The biological sciences
are concerned with fundamental questions about the nature of human beings and
our place in the natural universe--questions such as "What is life?" "How did
living organisms arise?" and "How are human beings related to and distinct from
other living things?" Common Core courses emphasize the core biological
concepts of continuity (genetics), organization, regulation, and evolution. The
biological sciences are concerned with many different levels of organization,
from cells and the molecules they produce to ecosystems and the species that
constitute them. To give students an appreciation of the range of subject
matter in the biological sciences, Common Core sequences are organized such
that one quarter emphasizes the biology of cells and molecules, one the biology
of organisms, and one the biology of populations or species. Students learn the
key concepts and methods to generate new knowledge in each of these areas. In
addition, students come to appreciate the interconnectedness of the biological
sciences and the importance of integrating different approaches to the study of
living things.
The Biological Sciences Concentration Program
The goals of the biological sciences concentration are to give students an
understanding of the currently accepted concepts in biology and the
experimental support for these concepts, and an appreciation of the gaps in our
current understanding and the opportunities for new research in this field. The
concentration is designed to prepare students for graduate or professional
study in the biological sciences and for careers in biology. As in the Common
Core, emphasis is placed on introducing students to the diversity of subject
matter and methods of investigation in the biological sciences. (NOTE: The
curriculum is currently under revision. For updated information, contact the
BSCD office.)General Chem 111-112-113 or 121-122-123
Education Math 131-132, 151-152, or 161-162
BioSci Common Core
3 Phys 121-122-123, 131-132-133, or 141-142-143
2 Chem 217-218 or 220-221
1 Stat 220
1 biochemistry (BioSci 200 or BchMB 301-302)
1 cell biology (BioSci 207)*
1 developmental biology (BioSci 221-229)*
1 organismal biology (BioSci 232-246, or 260)
14 (total)
Combined Bachelor of Arts/Master of Science Degree Program in Biochemistry and
Molecular Biology
This program is designed for those students who, early in their academic
careers, decide to pursue graduate study in biology at the molecular level. It
differs from the usual concentration in the biological sciences in that it
requires a central core of graduate courses in biochemistry and molecular
biology. These in turn require a background in physical chemistry.
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Faculty
KAN L. AGARWAL, Professor, Departments of Biochemistry & Molecular Biology
and Chemistry and the College